One generally overlooked "villain crossover" appears in the 1938 serial FLASH GORDON'S TRIP TO MARS.
Now, had Azura of the FLASH GORDON comic strip started out as a flunky to Ming, I would not deem any teamup between the two a crossover. But the original comics appearance of Azura, examined in this essay, shows that her realm was independent of Ming's, and even in the serial Azura seems her own woman, and not a henchperson. This may well be the only crossover in American serials, since most co-villains do not sport such independent charisma.
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